Definition
A perceptual phenomenon in which the observer recognizes portions of an object occluded by another object.
Introduction
In the three-dimensional world, objects that are farther from the observer are often partly occluded by other objects that are closer to the observer. Although the occluded portions of the farther objects cast no optical information on the observer’s retina, they are recognized as if the occluded portions are completed, a phenomenon called amodal completion (Michotte et al. 1991). In Fig. 1a, the observer may perceive an intact disk “behind” a square by completing the portion “hidden” by the square, although the retinal image is merely a Pac-Man-like shape (three-quarters of a circle) placed adjacent to the square, with no gap between them. (If there is a gap between them, the Pac-Man-like shape will be perceived as it is, as in Fig. 1b.)
References
Aust, U., & Huber, L. (2006). Does the use of natural stimuli facilitate amodal completion in pigeons? Perception, 35, 333–349.
Ban, H., Yamamoto, H., Hanakawa, T., Urayama, S., Aso, T., Fukuyama, H., & Ejima, Y. (2013). Topographic representation of an occluded object and the effects of spatiotemporal context in human early visual areas. Journal of Neuroscience, 33, 16992–17007.
Cerella, J. (1980). The pigeon’s analysis of pictures. Pattern Recognition, 12, 1–6.
Darmaillacq, A.-S., Dickel, L., Rahmani, N., & Shashar, N. (2011). Do reef fish, Variola louti and Scarus niger, perform amodal completion? Evidence from a field study. Journal of Comparative Psychology, 125, 273–277.
Deruelle, C., Barbet, I., Dépy, D., & Fagot, J. (2000). Perception of partly occluded figures by baboons (Papio papio). Perception, 29, 1483–1497.
Fagot, J., Barbet, I., Parron, C., & Deruelle, C. (2006). Amodal completion by baboons (Papio papio): Contribution of background depth cues. Primates, 47, 145–150.
Forkman, B. (1998). Hens use occlusion to judge depth in a two dimensional picture. Perception, 27, 861–867.
Fujita, K. (2001). Perceptual completion in rhesus monkeys (Macaca mulatta) and pigeons (Columba livia). Perception & Psychophysics, 63, 115–125.
Fujita, K., & Giersch, A. (2005). What perceptual rules do capuchin monkeys (Cebus Apella) follow in completing partly occluded figures? Journal of Experimental Psychology: Animal Behavior Processes, 31, 387–398.
Fujita, K., & Ushitani, T. (2005). Better living by not completing: A wonderful peculiarity of pigeon vision? Behavioural Processes, 69, 59–66.
Kanizsa, G., Renzi, P., Conte, S., Compostela, C., & Guerani, L. (1993). Amodal completion in mouse vision. Perception, 22, 713–721.
Kellman, P. J., & Shipley, T. F. (1991). A theory of visual interpolation in object perception. Cognitive Psychology, 23, 141–221.
Kellman, P. J., & Spelke, E. S. (1983). Perception of partly occluded objects in infancy. Cognitive Psychology, 15, 483–524.
Lea, S. E. G., Slater, A. M., & Ryan, C. M. E. (1996). Perception of object unity in chicks: A comparison with the human infant. Infant Behavior and Development, 19, 501–504.
Michotte, A., Thines, G., & Crabbe, G. (1991). Amodal completion of perceptual structures. In G. Thines, A. Costall, & G. Butterworth (Eds.), Michotte’s experimental phenomenology of perception (pp. 140–167). Hillsdale: Erlbaum. (Original work published in 1964).
Nagasaka, Y., Lazareva, O. F., & Wasserman, E. A. (2007). Prior experience affects amodal completion in pigeons. Perception & Psychophysics, 69, 596–605.
Nakamura, N., Watanabe, S., Betsuyaku, T., & Fujita, K. (2010). Do bantams (Gallus gallus domesticus) experience amodal completion? An analysis of visual search performance. Journal of Comparative Psychology, 124, 331–335.
Nakamura, N., Watanabe, S., Betsuyaku, T., & Fujita, K. (2011). Do bantams (Gallus gallus domesticus) amodally complete partly occluded lines? An analysis of line classification performance. Journal of Comparative Psychology, 125, 411–419.
Pepperberg, I. M., & Nakayama, K. (2016). Robust representation of shape in a Grey parrot (Psittacus erithacus). Cognition, 153, 146–160.
Rauschenberger, R., & Yantis, S. (2001). Masking unveils pre-amodal completion representation in visual search. Nature, 410, 369–372.
Regolin, L., & Vallortigara, G. (1995). Perception of partly occluded objects by young chicks. Perception & Psychophysics, 57, 971–976.
Sato, A., Kanazawa, S., & Fujita, K. (1997). Perception of object unity in a chimpanzee (Pan troglodytes). Japanese Psychological Research, 39, 191–199.
Sekuler, A. B. (1994). Local and global minima in visual completion effects of symmetry and orientation. Perception, 23, 529–545.
Sekuler, A. B., Lee, J. A. J., & Shettleworth, S. J. (1996). Pigeons do not complete partly occluded figures. Perception, 25, 1109–1120.
Shimizu, T. (1998). Conspecific recognition in pigeons (Columba livia) using dynamic video images. Behaviour, 135, 43–53.
Shipley, T. F., & Kellman, P. J. (1992). Perception of partly occluded objects and illusory figures: Evidence for an identity hypothesis. Journal of Experimental Psychology: Human Perception and Performance, 18, 106–120.
Sovrano, V. A., & Bisazza, A. (2008). Recognition of partly occluded objects by fish. Animal Cognition, 11, 161–166.
Sugita, Y. (1999). Grouping of image fragments in primary visual cortex. Nature, 401, 269–272.
Takahasi, M., & Okanoya, K. (2013). An invisible sign stimulus: Completion of occluded visual images in the Bengalese finch in an ecological context. Neuroreport, 24, 370–374.
Ushitani, T., & Fujita, K. (2005). Pigeons do not perceptually complete partly occluded photos of food: An ecological approach to the “pigeon problem”. Behavioural Processes, 69, 67–78.
Ushitani, T., Fujita, K., & Yamanaka, R. (2001). Do pigeons (Columba livia) perceive object unity? Animal Cognition, 4, 153–161.
Ushitani, T., Imura, T., & Tomonaga, M. (2010). Object-based attention in chimpanzees (Pan troglodytes). Vision Research, 50, 577–584.
Watanabe, S., & Furuya, I. (1997). Video display for study of avian visual cognition: From psychophysics to sign language. International Journal of Comparative Psychology, 10, 111–127.
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Ushitani, T. (2020). Amodal Completion. In: Vonk, J., Shackelford, T. (eds) Encyclopedia of Animal Cognition and Behavior. Springer, Cham. https://doi.org/10.1007/978-3-319-47829-6_1065-1
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DOI: https://doi.org/10.1007/978-3-319-47829-6_1065-1
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